Multivariate analysis of variance: Difference between revisions

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[[Image:Humpback underwater.jpg|thumb|300px|alt=A whale swims horizontally underwater, with long front flippers extended and with mottled skin.|[[Humpback whale]]s are well known for their songs{{Citation needed|date=July 2011}}]]
'''Whale sounds''' are used by [[whale]]s for different kinds of [[animal communication|communication]].<ref>Communication and behavior of whales, R Payne. 1983. Westview Press.</ref>
 
The mechanisms used to produce sound vary from one family of [[cetaceans]] to another. [[Marine mammal]]s, such as whales, [[dolphin]]s, and [[porpoise]]s, are much more dependent on sound for communication and sensation than are land mammals, because other senses are of limited effectiveness in water. [[Visual perception|Sight]] is less effective for marine mammals because of the way particulates in the ocean scatter [[light]]. [[Olfaction|Smell]] also is limited, as [[molecule]]s diffuse more slowly in water than in air, which makes smelling less effective. However, the [[speed of sound]] is roughly four times greater in water than in the atmosphere at [[sea level]]. Because sea mammals are so dependent on hearing to communicate and feed, environmentalists and [[cetology|cetologists]] are concerned that they are being harmed by the increased [[ambient noise]] in the world's oceans caused by [[ship]]s, [[sonar]] and marine seismic surveys.<ref name="Melcón2012"/>
 
The word "[[song]]" is used to describe the pattern of regular and predictable sounds made by some species of whales, notably the [[Humpback Whale]]. This is included with or in comparison with [[music]], and male humpback whales have been described as "inveterate [[composers]]" of songs that are "'strikingly similar' to human musical traditions".<ref>Payne Roger, quoted in: Author(s): Susan Milius. "Music without Borders", p. 253. Source: ''Science News'', Vol. 157, No. 16, (15 April 2000), pp. 252-254. Published by: Society for Science &amp; the Public.</ref> It has been suggested that humpback songs communicate male fitness to female whales.<ref>A.J. Wright and A Walsh (2010) Mind the gap: why neurological plasticity may explain seasonal interruption in humpback whale song. Journal of the Marine Biological Association of the United Kingdom, 90(8), 1489–1491.</ref> The click sounds made by [[Sperm whale]]s and dolphins are not strictly song, but the clicking sequences have been suggested to be individualized rhythmic sequences that communicate the identity of a single whale to other whales in its group and allows the groups to coordinate foraging activities.<ref>Michel Andre and Cees Kamminga (2000) Rhythmic dimension in the echolocation click trains of sperm whales: a possible function of identification and communication Journal of Marine Biological Association of the United Kingdom, Vol. 80, pp. 163-169.</ref>
 
==Production of sound==
Humans produce sound by expelling air through the [[larynx]]. The [[vocal cords]] within the larynx open and close as necessary to separate the stream of air into discrete pockets of air.<ref>{{cite web |url=http://www.dosits.org/animals/soundproduction/mammalsproduce/ |date=July 2009 |title=How do marine mammals produce sounds? |deadurl=no |accessdate=15 October 2013}}</ref> These pockets are shaped by the [[throat]], [[tongue]], and [[lip]]s into the desired sound, allowing humans to [[human voice|speak]].
 
Cetacean sound production differs markedly from this mechanism. The precise mechanism differs in the two major suborders of cetaceans:&nbsp;the&nbsp;''Odontoceti''&nbsp;([[toothed whales]]—including dolphins) and the ''Mysticeti'' ([[baleen whale]]s—including the largest whales, such as the [[Blue Whale]]).
 
===Odontocete whale sound production===
[[File:Delfinekko.gif|thumb|left|Process in a dolphin echolocation: in green the sounds generated by the dolphin, in red from the fish.]]
[[Image:Delphinschaedel.svg|300px|thumb|alt=Outline of what's inside a dolphin head. The skull is to the rear of the head, with the jaw bones extending narrowly forward to the nose. The anterior bursa occupies most of the upper front of the head, ahead of the skull and above the jaw. A network of air passages run from the upper roof of the mouth, past the back of the anterior bursa, to the [[blowhole]]. The posterior bursa is a small region behind the air passages, opposite the anterior bursa. Small phonic tips connect the bursa regions to the air passages.|Idealized dolphin head showing the regions involved in sound production. This image was redrawn from Cranford (2000).]]
Odontocetes produce rapid bursts of high-frequency clicks that are thought to be primarily for [[animal echolocation|echolocation]]. Specialized organs in an odontocete produce collections of clicks and buzzes at frequencies from 0.2 to 150&nbsp;kHz to obtain sonic information about its environment. Lower frequencies are used for distance echolocation, due to the fact that shorter wavelengths do not travel as far as longer wavelengths underwater. Higher frequencies are more effective at shorter distances, and can reveal more detailed information about a target. Echoes from clicks convey, not only the distance to the target, but also the size, shape, speed, and vector of its movement. Additionally, echolocation allows the odontocete to easily discern the difference between objects that are different in material composition, even if visually identical, by their different densities. Individuals appear to also be able to isolate their own echoes during pod feeding activity without interference from other pod members' echolocations.<ref name="dolphins.org">http://www.dolphins.org/marineed_acoustics.php</ref>
 
Whistles are used for communication, and four- to six-month-old calves develop unique sounds that they use most frequently throughout their lives. Such "signature whistles" are distinctive to the individual and may serve as a form of identification among other odontocetes.<ref name="dolphins.org"/> Though a large pod of dolphins will produce a wide range of different noises, very little is known about the meaning of the sound. Frankel quotes one researcher who says listening to a school of odontocetes is like listening to a group of children at a school playground.<ref name=Frankel>Frankel, Adam S. "Sound production", ''Encyclopedia of Marine Mammals'', 1998, pp. 1126–1137. ISBN 0-12-551340-2.</ref>
 
The multiple sounds odontocetes make are produced by passing air through a structure in the head called the '''phonic lips'''.{{Citation needed|date=July 2009}} This structure functions like the human nasal cavity.{{Citation needed|date=July 2009}} As the air passes through this narrow passage, the phonic lip membranes are sucked together, causing the surrounding tissue to vibrate. These vibrations can, as with the vibrations in the human larynx, be consciously controlled with great sensitivity.{{Citation needed|date=July 2009}} The vibrations pass through the tissue of the head to the [[melon (whale)|melon]], which shapes and directs the sound into a beam of sound useful in echolocation. Every toothed whale except the [[sperm whale]] has two sets of phonic lips and is thus capable of making two sounds independently.<ref>{{cite journal | author = Fitch | coauthors = W.T. Neubauer; and Herzel J. | year = 2002 | title = Calls out of chaos: the adaptive significance of nonlinear phenomena in mammalian vocal production| journal = Anim. Behav. | volume = 63 | pages = 407–418 | doi = 10.1006/anbe.2001.1912 | issue = 3 }}</ref> Once the air has passed the phonic lips it enters the [[vestibular system|vestibular sac]]. From there, the air may be recycled back into the lower part of the nasal complex, ready to be used for sound creation again, or passed out through the blowhole.{{Citation needed|date=July 2009}}
 
The [[French language|French]] name for phonic lips, ''museau de singe'', translates literally as "monkey's muzzle," which the phonic lip structure is supposed to resemble.<ref>{{cite web |url=http://www.spermwhale.org/SpermWhale/spermwhaleorgV1.html |title=Selected Whale Sciences Images - Volume 1 |author=Ted W. Cranford |accessdate=20 October 2010}}</ref> New cranial analysis using [[computed axial tomography|computed axial]] and [[single photon emission computed tomography]] scans in 2004 showed, at least in the case of [[bottlenose dolphin]]s, that air might be supplied to the nasal complex from the lungs by the palatopharyngeal sphincter, enabling the sound creation process to continue for as long as the dolphin is able to hold its breath.<ref>{{cite journal | author = Houser, Dorian S. | coauthors = Finneran, James; Carder, Don; Van Bonn, William; Smith, Cynthia; Hoh, Carl; Mattrey, Robert; and Ridgway, Sam | year = 2004 | title = Structural and functional imaging of bottlenose dolphin (''Tursiops truncatus'') cranial anatomy | journal = Journal of Experimental Biology | volume = 207 | pages = 3657–3665 | doi = 10.1242/jeb.01207 | pmid = 15371474 | issue = Pt 21}}</ref>
 
===Mysticete whale sound production===
[[Baleen whale]]s (formally called mysticetes) do not have phonic lip structure. Instead, they have a larynx that appears to play a role in sound production, but it lacks vocal cords, and scientists remain uncertain as to the exact mechanism.{{Citation needed|date=July 2009}} The process, however, cannot be completely analogous to humans, because whales do not have to exhale in order to produce sound. It is likely that they recycle air around the body for this purpose. Cranial sinuses may also be used to create the sounds, but again researchers are currently unclear how.
 
===Mysticete whale sound levels===
The frequency of baleen whale sounds ranges from 10 [[Hertz|Hz]] to 31&nbsp;kHz.<ref name=RGMT>{{cite book |title=Marine Mammals and Noise|author=Richardson, Greene, Malme, Thomson|year=1995|publisher=Academic Press|isbn=978-0-12-588440-2}}</ref> A list of typical levels is shown in the table below.
 
{| class="wikitable"
|-
! Source
! [[Broadband]] source level (dB re 1 <math>\mu</math>Pa at 1m)<ref name=KupermanRoux>{{cite book |author=Kuperman, Roux| editor=Rossing, Thomas D. |title=Springer Handbook of Acoustics |publisher=Springer |year=2007 | chapter = Underwater Acoustics |isbn=978-0-387-30446-5}}</ref>
|-
| [[Fin whale]] moans
| 155–186
|-
| [[Blue whale]] moans
| 155–188
|-
| [[Gray whale]] moans
| 142–185
|-
| [[Bowhead whale]] tonals, moans and song
| 128–189
|}
 
==Purpose of whale-created sounds==
While the complex sounds of the Humpback whale (and some blue whales) are believed to be primarily used in [[sexual selection]],{{Citation needed|date=October 2013}} <!-- Is the use of the term "sexual selection" correct here, and below in the next section?--> the simpler sounds of other whales have a year-round use.{{Citation needed|date=July 2009}} While toothed whales are capable of using echolocation to detect the size and nature of objects, this capability has never been demonstrated in baleen whales.{{Citation needed|date=July 2009}} Further, unlike some fish such as [[shark]]s, a whale's sense of smell is not highly developed.{{Citation needed|date=September 2012}} Thus, given the poor visibility of aquatic environments and that sound travels so well in water, sounds audible to humans may play a role in navigation. For instance, the depth of water or the existence of a large obstruction ahead may be detected by loud noises made by baleen whales.{{Citation needed|date=July 2009}}
 
The singing of whale songs for [[aesthetics|aesthetic]] enjoyment, personal satisfaction, or 'for art's sake', is "an untestable question in scientific terms."<ref>Entomologist Thomas Eisner, quoted in: Milius (2000), p. 254 though the production of sound out of ecstasy, is common among the toothed species.</ref>
 
==Song of the Humpback Whale==
[[Image:HumBack2.jpg|right|thumb|300px|[[Spectrogram]] of humpback whale vocalizations. Detail is shown for the first 24 seconds of the 37 second recording humpback whale song. The ethereal whale songs and echolocation clicks are visible as horizontal striations and vertical sweeps respectively.{{citation needed|date=December 2012}}]] {{Listen|filename=Humpbackwhale2.ogg|title=Humpback Whale Song|description=Recording of humpback whales singing and clicking.|format=[[Ogg]]}}
 
Two groups of whales, the humpback whale and the subspecies of blue whale found in the [[Indian Ocean]], are known to produce a series of repetitious sounds at varying frequencies known as whale song. Marine biologist Philip Clapham describes the song as "probably the most complex in the animal kingdom."<ref>{{cite book | title = Humpback whales | author = Clapham, Philip | publisher = Colin Baxter Photography | year = 1996 | isbn = 0-948661-87-9}}</ref>
 
Male humpback whales perform these vocalizations often during the mating season, and so it is believed the purpose of songs is to aid mate selection.<ref name=Frankel/> Whether the songs are a competitive behavior between males seeking the same mate, a means of defining territory or a "flirting" behavior from a male to a female is not known and the subject of ongoing research.{{Citation needed|date=July 2009}} Males have been observed singing while simultaneously acting as an escort whale in the immediate vicinity of a female. Singing has also been recorded in competitive groups of whales that are composed of one female and multiple males.{{Citation needed|date=July 2009}}
 
Interest in whale song was aroused by researchers [[Roger Payne]] and [[Scott McVay]] after the songs were brought to their attention by a Bermudian named Frank Watlington who was working for the US government at the SOFAR station listening for Russian submarines with underwater hydrophones off the coast of the island.<ref>{{cite book | title = Thousand mile song | author = Rothenberg, David | publisher = Basic Books | year = 2008 | isbn = 978-0-465-07128-9}}</ref>
 
The songs follow a distinct hierarchical structure. The base units of the song (sometimes loosely called the "[[note (music)|notes]]") are single uninterrupted emissions of sound that last up to a few seconds. These sounds vary in frequency from 20&nbsp;Hz to upward of 24&nbsp;kHz (the typical human range of hearing is 20&nbsp;Hz to 20&nbsp;kHz). The units may be [[frequency modulation|frequency modulated]] (i.e., the pitch of the sound may go up, down, or stay the same during the note) or [[amplitude modulation|amplitude modulated]] (get louder or quieter). However the adjustment of bandwidth on a spectrogram representation of the song reveals the essentially [[Pulse (signal processing)|pulsed]] nature of the FM sounds.
 
A collection of four or six units is known as a sub-[[phrase]], lasting perhaps ten seconds (see also [[phrase (music)]]).{{Citation needed|date=July 2009}} A collection of two sub-phrases is a phrase. A whale will typically repeat the same phrase over and over for two to four minutes. This is known as a theme. A collection of themes is known as a song.{{Citation needed|date=July 2009}} The whale will repeat the same song, which last up to 30 or so minutes, over and over again over the course of hours or even days.{{Citation needed|date=July 2009}} This "[[Russian doll]]" hierarchy of sounds has captured the imagination of scientists.{{Weasel-inline|date=July 2009}}{{Citation needed|date=July 2009}}
 
All the whales in an area sing virtually the same song at any point in time and the song is constantly and slowly evolving over time.{{Citation needed|date=July 2009}} For example, over the course of a month a particular unit that started as an upsweep (increasing in frequency) might slowly flatten to become a constant note.<ref name=Frankel/> Another unit may get steadily louder. The pace of evolution of a whale's song also changes—some years the song may change quite rapidly, whereas in other years little variation may be recorded.<ref name=Frankel/>
 
{{Wide image|Humpback song.PNG|1200px|Idealized schematic of the song of a humpback whale.<br/>Redrawn from Payne, et al. (1983)|600px|center|alt=Six long parallel lines with tick marks. "Song session (hours–days)" has no ticks. "Song (12–15 mins)" has 1 tick. "Theme (2 mins)" has 4 ticks. "Phrase (15–20 secs)" has 18 ticks. "Sub-phrase (7 secs)" has 36 ticks. "Unit (1 sec)" has many more ticks, this time angled up or down; it also has many gaps in the line.}}
 
[[File:Akhumps 128 016 0 500c.gif|left|thumb|255px|alt=Two spectral images with X axis being time. In one, the Y axis is frequency and there is a complicated pattern in the 10–450 Hz region. In the other, the Y axis is amplitude, which is largely constant but with many small spikes.|Humpback whale, sound spectrum and time plots]]Whales occupying the same geographical areas (which can be as large as entire ocean basins) tend to sing similar songs, with only slight variations. Whales from non-overlapping regions sing entirely different songs.<ref name=Frankel/>
 
As the song evolves, it appears that old patterns are not revisited.<ref name=Frankel/> An analysis of 19 years of whale songs found that while general patterns in song could be spotted, the same combination never recurred.{{Citation needed|date=July 2009}}
 
Humpback whales may also make stand-alone sounds that do not form part of a song, particularly during courtship rituals.<ref name=JMamm>{{cite journal|doi=10.2307/1381574|jstor=1381574|title=Humpback Whale Songs on a North Atlantic Feeding Ground|author=Mattila, David. K|coauthors=Guinee, Linda N., Mayo, Charles A.|publisher=American Society of Mammalogists|year=1987|volume=68|issue=4 |pages=880–883|journal=Journal of Mammalogy |url=http://www.jstor.org/discover/10.2307/1381574?uid=3739256&uid=2&uid=4&sid=21102778038323 |deadurl=no |accessdate=15 October 2013}}</ref> Finally, humpbacks make a third class of sound called the feeding call.{{Citation needed|date=July 2009}} This is a long sound (5 to 10 s duration) of near constant frequency. Humpbacks generally feed cooperatively by gathering in groups, swimming underneath shoals of fish and all lunging up vertically through the fish and out of the water together. Prior to these lunges, whales make their feeding call. The exact purpose of the call is not known, but research suggests that fish know what it means.{{Citation needed|date=July 2009}} When the sound was played back to them, a group of herring responded to the sound by moving away from the call, even though no whale was present.{{Citation needed|date=July 2009}}
 
Some scientists have proposed that humpback whale songs may serve an [[animal echolocation|echolocative]] purpose,<ref>{{cite journal |author=Mercado, E. III, and Frazer, L.N. |year=2001 |title=Humpback whale song or humpback whale sonar? A Reply to Au ''et al.'' |journal=IEEE Journal of Oceanic Engineering |volume=26 |pages=406–415 |url=http://www.acsu.buffalo.edu/~emiii/00946514.pdf |format=PDF |doi=10.1109/48.946514 |issue = 3}}</ref> but this has been subject to disagreement.<ref>{{cite journal | author = W. W. L. Au, A. Frankel, D. A. Helweg, and D. H. Cato |year=2001 |title=Against the humpback whale sonar hypothesis | journal = IEEE Journal of Oceanic Engineering | volume = 26 | pages = 295–300 | doi = 10.1109/48.922795 | issue = 2}}</ref>
 
==Other whale sounds==
 
Humpback whales have also been found to make a range of other social sounds to communicate such as "grunts", "groans", "thwops", "snorts" and "barks".<ref>Cecilia Burke, '[http://www.australiangeographic.com.au/journal/a-whales-varied-vocabulary.htm 'A whale's varied vocabulary', Australian Geographic], AG Online. Retrieved 7 August 2010.</ref>
 
Most baleen whales make sounds at about 15–20 [[hertz]].{{Citation needed|date=July 2009}} However, a team of [[Marine biology|marine biologists]], led by Mary Ann Daher of the [[Woods Hole Oceanographic Institution]], reported in ''[[New Scientist]]'' in December 2004 that they had been tracking a whale in the North Pacific for 12 years that was "singing" at 52&nbsp;Hz.<ref name=NewSci>{{cite news|url=http://www.newscientist.com/article/mg18424774.600-lonely-whales-song-remains-a-mystery.html|date=11 December 2004|accessdate=12 July 2009|title=Lonely whale's song remains a mystery|publisher=[[Reed Business Information Ltd]]|work=[[New Scientist]]}}</ref> The scientists are unable to explain this dramatic difference from the norm; however, they believe the whale is baleen<ref>{{cite news|url=http://www.dailytimes.com.pk/default.asp?page=story_13-12-2004_pg6_14|title=Strange-voiced whale at large in the ocean|author=Associated Press|date=13 December 2004|accessdate=12 July 2009|publisher=''[[Daily Times (Pakistan)|Daily Times]]''}}</ref> and unlikely to be a new species,<ref name=NewSci/> suggesting that currently known species may have a wider vocal range than previously thought.
 
In 2009, researchers found that [[blue whale]] song has been deepening in its tonal frequency since the 1960s.<ref>McDonald, Mark A., Hildebrand, John A., Mesnick, Sarah. [http://www.int-res.com/abstracts/esr/v9/n1/p13-21/ Worldwide decline in tonal frequencies of blue whale songs.] ''Endangered Species Research, Vol. 9 No. 1'' 23 October 2009.</ref> While [[noise pollution]] has increased ambient ocean noise by over 12 decibels since the mid-20th century, researcher Mark McDonald indicated that higher pitches would be expected if the whales were straining to be heard.<ref>Keim, Brandon. [http://www.wired.com/wiredscience/2009/12/blue-whale-song-mystery/ Blue Whale Song Mystery Baffles Scientists.] ''[[Wired (magazine)]].'' 2 December 2009.</ref>
 
Most other whales and dolphins produce sounds of varying degrees of complexity. Of particular interest is the [[Beluga (whale)|Beluga]] (the "sea canary") which produces an immense variety of whistles, clicks and pulses.{{Citation needed|date=July 2009}}
 
==Human interaction==
[[Image:Example of Blue Whales' D calls in presence of MFA sonar - Melcón et al. 2012.png|thumb|right|Blue whales stop producing foraging D calls once a mid-frequency sonar is activated, even though the sonar frequency range (1–8 kHz) far exceeds their sound production range (25–100 Hz).<ref name="Melcón2012"/>]]
[[Image:The Sounds of Earth - GPN-2000-001976.jpg|right|thumb|alt=Flat circular disc of gold, with a central label, a hole, and a wide band of very small lines, like a golden version of an old analog record|[[Voyager Golden Record]]s carried whale songs into [[outer space]] with other sounds representing planet Earth.]]
Though some observers suggest that undue fascination has been placed on the whales' songs simply because the animals are under the sea,{{Citation needed|date=July 2009}} most marine mammal scientists believe that sound plays a particularly vital role in the development and well-being of cetaceans.{{Citation needed|date=July 2009}}
 
Researchers use [[hydrophone]]s (often adapted from their original military use in tracking submarines) to ascertain the exact location of the origin of whale noises.{{Citation needed|date=July 2009}} Their methods allow them also to detect how far through an ocean a sound travels.{{Citation needed|date=September 2012}} Research by Dr. Christopher Clark of [[Cornell University]] conducted using military data showed that whale noises travel for thousands of kilometres.<ref name=BBC>{{cite news|url=http://news.bbc.co.uk/1/hi/sci/tech/4297531.stm|author=Bentley, Molly|date=28 February 2005|accessdate=12 July 2009 | work=BBC News | title=Unweaving the song of whales}}</ref> As well as providing information about song production, the data allows researchers to follow the migratory path of whales throughout the "singing" (mating) season. An important finding is that whales, in a process called the [[Lombard effect]], adjust their song to compensate for background [[noise pollution]].<ref name=" Scheifele">{{Cite journal|pmid=15807036|year=2005|last1=Scheifele|first1=PM|last2=Andrew|first2=S|last3=Cooper|first3=RA|last4=Darre|first4=M|last5=Musiek|first5=FE|last6=Max|first6=L|title=Indication of a Lombard vocal response in the St. Lawrence River Beluga|volume=117|issue=3 Pt 1|pages=1486–92|journal=The Journal of the Acoustical Society of America}}</ref> Moreover, there is evidence that blue whales stop producing foraging D calls once a mid-frequency sonar is activated, even though the sonar frequency range (1–8&nbsp;kHz) far exceeds their sound production range (25–100&nbsp;Hz).<ref name="Melcón2012">{{Cite journal|doi=10.1371/journal.pone.0032681|title=Blue Whales Respond to Anthropogenic Noise|year=2012|editor1-last=Mathevon|editor1-first=Nicolas|last1=Melcón|first1=Mariana L.|last2=Cummins|first2=Amanda J.|last3=Kerosky|first3=Sara M.|last4=Roche|first4=Lauren K.|last5=Wiggins|first5=Sean M.|last6=Hildebrand|first6=John A.|journal=PLoS ONE|volume=7|issue=2|pages=e32681|pmid=22393434|pmc=3290562}}</ref>
 
Prior to the introduction of human noise production, Clark says the noises may have travelled right from one side of an ocean to the other, agreeing with a thirty-year-old concept blaming large-scale shipping.<ref name=BBC/> His research indicates that ambient noise from boats is doubling with each decade.<ref name=BBC/> This has the effect of reducing the range at which whale noises can be heard. [[Environmentalist]]s fear that such boat activity is putting undue stress on the animals as well as making it difficult to find a mate.<ref name=BBC/>
 
In the past decade, many effective automated methods, such as Signal Processing, Data Mining and Machine Learning techniques have been developed to detect and classify the Whale vocalization.<ref> M. Pourhomayoun, P. Dugan, M. Popescu, and C. Clark, “Bioacoustic Signal Classification Based on Continuous Region Features, Grid Masking Features and Artificial Neural Network,” International Conference on Machine Learning (ICML), 2013.</ref><ref>7. M. Popescu, P. Dugan, M. Pourhomayoun, and C. Clark, “Periodic Pulse Train Signal Detection and Classification using Spectrogram Intensity Binarization and Energy Projection,” International Conference on Machine Learning (ICML), 2013.</ref>
 
==Media==
{{See also|List of whale vocalizations}}
{{listen
| filename    = Humpback whale wheezeblow.ogg
| title        = A Humpback Whale song
| description  =
| format      = [[Ogg]]
| filename2    = Humpback whale moo.ogg
| title2      = Common Humpback Whale vocalizations on a windy day
| description2 =
| format2      = [[Ogg]]
| filename3    = Killer whale.ogg
| title3      = An Orca's song
| description3 =
| format3      = [[Ogg]]
| filename4    = Killer whale simple.ogg
| title4      = An Orca's song, at a distance
| description4 =
| format4      = [[Ogg]]
| filename5    = Killer whale residents broadband.ogg
| title5      = Vocalizations of an Orca
| description5 =
| format5      = [[Ogg]]
}}
 
==Selected discography==
 
* ''Songs of the Humpback Whale'' (SWR 118) was originally released in 1970 by [[CRM Records]] from recordings made by [[Roger Payne]], Frank Watlington, and others. The LP was later re-released by [[Capitol Records]], published in a flexible format in the National Geographic Society magazine, Volume 155, Number 1, in January 1979, re-released by Living Music/Windham Hill/BMG Records on CD in 1992, and remastered on CD by BGO-Beat Goes On in 2001.
 
* ''Deep Voices: The Second Whale Record'' (Capitol/EMI Records 0777 7 11598 1 0) was released on LP in 1977 from additional recordings made by [[Roger Payne]], and re-released on CD in 1995 by Living Music/Windham Hill/BMG Records. It includes recordings of humpbacks, blues, and rights.
 
* ''Northern Whales'' (MGE 19) was released by [[Music Gallery Editions]] from recordings made by Pierre Ouellet, John Ford, and others affiliated with [[Interspecies Music and Communication Research]]. It includes recordings of belugas, narwhals, orca, and bearded seals.
 
* ''Sounds of the Earth: Humpback Whales'' (Oreade Music) was released on CD in 1999.
 
* ''Rapture of the Deep: Humpback Whale Singing'' (Compass Recordings) was released on CD in 2001.
*[http://www.songlinesofthewhales.org/ Songlines: Songs of the East Australian Humpback whales.] was released in 2009.
 
==See also==
* [[52-Hertz whale]]
*[[Bioacoustics]]
*[[Biomusic]]
*[[Underwater acoustics]]
*[[Vocal learning]]
{{Portal|left=yes|Cetaceans}}
 
==References==
{{Reflist|2}}
 
===General references===
 
<div class="references-small">
*''Lone whale's song remains a mystery'', ''New Scientist'', issue number 2477, 11 December 2004
* {{cite journal
| author = Frazer, L.N. and Mercado. E. III.
| year = 2000
| title = A sonar model for humpback whale song
| journal = IEEE Journal of Oceanic Engineering
| volume = 25 | pages = 160–182
| doi = 10.1109/48.820748}}
* Helweg, D.A., Frankel, A.S., Mobley Jr, J.R. and [[Louis Herman|Herman, L.M.]], “Humpback whale song: our current understanding,” in ''Marine Mammal Sensory Systems'', J. A. Thomas, R. A. Kastelein, and A. Y. Supin, Eds. New York: Plenum, 1992, pp.&nbsp;459–483.
*''In search of impulse sound sources in odontocetes'' by Ted Cranford in ''Hearing by whales and dolphins'' (W. Lu, A. Popper and R. Fays eds.). Springer-Verlag (2000).
*''Progressive changes in the songs of humpback whales (''Megaptera novaeangliae''): a detailed analysis of two seasons in Hawaii'' by K.B.Payne, P. Tyack and R.S. Payne in ''Communication and behavior of whales''. Westview Press (1983)
* {{cite news | publisher=BBC News | date= 28 February 2005| url=http://news.bbc.co.uk/1/hi/sci/tech/4297531.stm
| title=Unweaving the song of whales}}
</div>
 
==External links==
{{Commons category|Audio files of Balaenopteridae}}
*[http://www.voicesinthesea.org/ Voices in the Sea] has Whale and Dolphin sounds and interpretive videos
*[http://www.birds.cornell.edu/brp/ Cornell University's Bioacoustics Research Program]
*[http://www.listenforwhales.org/ Right Whale Listening Network], a project of the above bioacoustics program at Cornell
*[http://www.avantgardeproject.org/AGP28/index.htm Whale Songs at the Avant Garde Project] has FLAC files made from high-quality LP transcriptions.
*[https://www.youtube.com/watch?v=2DoX-zOQok0 Perspectives in Ocean Science] Listening to Whales, John Hildebrand, Scripps Institution of Oceanography
*[http://www.bl.uk/soundarchive The British Library Sound Archive] contains over 150,000 recordings of animal sounds and natural atmospheres from around the world.
*[http://www.songlinesofthewhales.org/ Songlines: Songs of the East Australian Humpback whales.]
*Recording of the bearded seal's "spiralling trill," one of the most phenomenal vocalizations of the underwater kindgdom: http://www.birds.cornell.edu/brp/listen-to-project-sounds/bearded-seal
 
{{Animal communication}}
{{Hydroacoustics}}
{{Zoomusicology}}
 
{{DEFAULTSORT:Whale Song}}
[[Category:Animal intelligence]]
[[Category:Song forms]]
[[Category:Whale sounds]]
 
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{{Link FA|ca}}
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Latest revision as of 19:42, 20 December 2014

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